Revisions to Vintana and Adalatherium move them to the stylinodontids
This post repairs earlier posts
as part of the housekeeping process. No one presented the following hypothesis of interrelationships in prior publications. This hypothesis now requires confirmation, refutation or modification with a similar taxon list.
Earlier
tiny Groeberia moved to the stylinodontids, leaving Vintana (Fig 1) and Adalatherium (Fig 2) behind somewhere in the Marsupialia, not far from diprotodontids.
Figure 1. Vintana revised with new DGS colors.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/vintana588-1.jpg?w=294″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/vintana588-1.jpg?w=584″ class=”size-full wp-image-91192″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/vintana588-1.jpg” alt=”Figure 1. Vintana revised with new DGS colors.” width=”584″ height=”596″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/vintana588-1.jpg?w=584&h=596 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/vintana588-1.jpg?w=147&h=150 147w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/vintana588-1.jpg?w=294&h=300 294w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/vintana588-1.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 1. Vintana revised with new DGS colors.
New interpretations and scoring
for Vintana (Fig 1) and Adalatherium (Fig 2) reunite these two with Groeberia in the stylinodontids (Fig 3), a clade between Primates and Phenacodontids and the rest of the large hooved herbivores.
Figure 2. Adalatherium revised with new DGS colors.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/adalatherium_skull588.jpg?w=157″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/adalatherium_skull588.jpg?w=534″ class=”size-full wp-image-91194″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/adalatherium_skull588.jpg” alt=”Figure 2. Adalatherium revised with new DGS colors.” width=”584″ height=”1119″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/adalatherium_skull588.jpg?w=584&h=1119 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/adalatherium_skull588.jpg?w=78&h=150 78w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/adalatherium_skull588.jpg?w=157&h=300 157w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/adalatherium_skull588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 2. Adalatherium revised with new DGS colors.
This clade is unusual in many regards,
one includes the upper canines anterior to the lower canines. In derived taxa the lower incisors form a virtual predentary.
Figure 1. Agriochoerus latifrons is now basal to Lophiodon and the stylinodontids in the LRT.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/agriochoerus-stylinodon588-2.jpg?w=66″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/agriochoerus-stylinodon588-2.jpg?w=226″ class=”size-full wp-image-91177″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/agriochoerus-stylinodon588-2.jpg” alt=”Figure 1. Agriochoerus latifrons is now basal to Lophiodon and the stylinodontids in the LRT.” width=”584″ height=”2649″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/agriochoerus-stylinodon588-2.jpg?w=584&h=2649 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/agriochoerus-stylinodon588-2.jpg?w=33&h=150 33w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/agriochoerus-stylinodon588-2.jpg?w=66&h=300 66w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/agriochoerus-stylinodon588-2.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 1. Agriochoerus latifrons is now basal to Lophiodon and the stylinodontids in the LRT.
Key thought worth considering:
The Late Cretaceous chronology of Vintana and Adalatherium means all more primitive taxa were also radiating in the Late Cretaceous. Notharctus was the only arboreal taxon. Vintana and Adalatherium were terrestrial, apparently surviving and thriving side-by-side with theropod dinosaurs.
Adalatherium hui
(Krause et al. 2020; Late Cretaceous) was originally considered a relative of multituberculates. The LRT nests Adalatherium with stylinodontid adapids within the clade Primates. The large adapid also from Madagascar, Megaladapis is an ancestor.
Vintana sertichi
(Krause et al. 2014; Late Cretaceous, UA 9972; 12.4 cm skull length; is a Madagascar mammal originally considered a member of the Allotheria and Gondwanatheria, two clades that do not appear in the large reptile tree. The LRT nests Vintana with stylinodontid adapids within the clade Primates. The large adapid also from Madagascar, Megaladapis is an ancestor.
Krause et al 2014 reported,
“The new taxon is the largest known mammaliaform from the Mesozoic of Gondwana. Its craniofacial anatomy reveals that it was herbivorous, large-eyed and agile, with well-developed high-frequency hearing and a keen sense of smell. The cranium exhibits a mosaic of primitive and derived features, the disparity of which is extreme and probably reflective of a long evolutionary history in geographic isolation.”
References
Krause DW, Hoffmann S, Wible JR, Kirk EC, and several other authors 2014. First cranial remains of a gondwanatherian mammal reveal remarkable mosaicism. Nature. online. doi:10.1038/nature13922. ISSN 1476-4687.
Krause DW et al. 2014. Vintana sertichi (Mammalia, Gondwanatheria) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology Memoir 14. 222pp.
Krause DW et al (10+ co-authors) 2020. Skeleton of a Cretaceous mammal from Madagascar reflects long-term insularity. Nature. 581(7809): 421–427.
Krause DW et al 2020b. Craniofacial morphology of Adalatherium hui (Mammalia, Gondwanatheria) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology. 40 (sup1): 19–66.
Source: https://pterosaurheresies.wordpress.com/2025/01/18/revisions-to-vintana-and-adalatherium-move-them-to-the-stylinodontids/
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