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Thylacoleo ancestors in the LRT

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Short one today
as graphics (Fig 1) tell the tale.

Figure 1. Thylacoleo ancestors in the LRT include Wakaleo, the Apatemyidae + Labidolemur. ” data-image-caption=”

Figure 1. Thylacoleo ancestors in the LRT include Wakaleo, the Apatemyidae + Labidolemur.

” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/dactylopsila.labidolemur588-1.jpg?w=75″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/dactylopsila.labidolemur588-1.jpg?w=257″ class=”size-full wp-image-91217″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/dactylopsila.labidolemur588-1.jpg” alt=”Figure 1. Thylacoleo ancestors in the LRT include Wakaleo, the Apatemyidae + Labidolemur.” width=”584″ height=”2326″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/dactylopsila.labidolemur588-1.jpg?w=584&h=2326 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/dactylopsila.labidolemur588-1.jpg?w=38&h=150 38w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/dactylopsila.labidolemur588-1.jpg?w=75&h=300 75w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/dactylopsila.labidolemur588-1.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />

Figure 1. Thylacoleo ancestors in the LRT include Wakaleo, the Apatemyidae + Labidolemur.

Once again
phylogenetic miniaturization from Dactylopsila to Labidolemur (Fig 1) marks the genesis of the Apatemyidae, now including the marsupial lions, Thylacoleo and Wakaleo (Fig 1) as their largest members.

We first looked at Wakaleo and Apatemys a few weeks ago here.

Figure 1. The extant striped opossum Dactylopsila in vivo. ” data-image-caption=”

Figure 1. The extant striped opossum Dactylopsila in vivo.

” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/07/dactylopsila_trivirgata588invivo.jpg?w=279″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/07/dactylopsila_trivirgata588invivo.jpg?w=584″ class=”size-full wp-image-87375″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/07/dactylopsila_trivirgata588invivo.jpg” alt=”Figure 1. The extant striped opossum Dactylopsila in vivo.” width=”584″ height=”628″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/07/dactylopsila_trivirgata588invivo.jpg?w=584&h=628 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/07/dactylopsila_trivirgata588invivo.jpg?w=140&h=150 140w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/07/dactylopsila_trivirgata588invivo.jpg?w=279&h=300 279w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/07/dactylopsila_trivirgata588invivo.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />

Figure 2. The extant striped opossum Dactylopsila in vivo. This is an extant late-surviving ancestor to Thylacoleo in the LRT.

Extant Dactylopsila
(Fig 2) was basal to several marsupial clades in the LRT.

PS
At the same time a tiny Late Jurassic echidna-like monotreme, Fruitafossor, has taken a second-tier (red) position in the LRT. It ws removed from the matrix for lacking any skull parts. A mandible is known. That brought about more resolution to the LRT.

This still appears to be a novel hypothesis of interrelationships.
If not, please provide a citation so I can promote it here.

References
Gillespie AK 2023. Two new marsupial lion taxa (Marsupialia, Thylacoleonidae) from the early and Middle Miocene of Australia. Alcheringa: An Australasian Journal of Palaeontology: 1–16.

wiki/Apatemyidae
wiki/Thylacoleo

Apatemyids and marsupial ‘lions’ now nest together in the LRT


Source: https://pterosaurheresies.wordpress.com/2025/01/19/thylacoleo-ancestors-in-the-lrt/


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